Kevin,
The original grips (1918) were numbered to the gun with the last two digits. I don't remember exactly but I think that the E/655 showed up around 1940-41. So I think the grips were cut to fit over an already clipped mag safety. The E stamped on the grip is an inspectors mark. You will also find initials or numbers randomly stamped on other parts of the gun. They were applied at different stages of production.
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Mike C.
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